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Petr Smýkal, Oldřich Trněný, Jan Brus, Pavel Hanáček, Abhishek Rathore, Rani Das Roma, Vilém Pechanec, Martin Duchoslav, Debjyoti Bhattacharyya, Michalis Bariotakis, Stergios Pirintsos, Jens Berger, Cengiz Toker
Genetic structure of wild pea (Pisum sativum subsp. elatius) populations in the northern part of the Fertile Crescent reflects moderate cross-pollination and strong effect of geographic but not environmental distance
www.plosone.org
www.plosone.org
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(to)Tj
1.0205 0 Td
(22)Tj
1.1735 0 Td
(individuals)Tj
4.6488 0 Td
(per)Tj
1.5307 0 Td
(population\))Tj
4.9323 0 Td
(of)Tj
1.0148 0 Td
(wild)Tj
1.9445 0 Td
(pea)Tj
1.5931 0 Td
(\()Tj
/F14 1 Tf
[()]TJ
/F8 1 Tf
(.)Tj
/F14 1 Tf
1.3379 0 Td
[()-3()22()7()15()-13()12()]TJ
/F8 1 Tf
-33.5109 -1.2982 Td
(subsp.)Tj
/F14 1 Tf
2.7155 0 Td
[( )15(\n)4()22()7()12()-4()]TJ
/F8 1 Tf
(\))Tj
3.1011 0 Td
(in)Tj
1.0318 0 Td
(the)Tj
1.474 0 Td
(region)Tj
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5.6523 0 Td
(Turkey.)Tj
3.3278 0 Td
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1.6101 0 Td
(consider)Tj
3.651 0 Td
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(representatives)Tj
6.1964 0 Td
(of)Tj
1.0148 0 Td
(pea)Tj
/F14 1 Tf
-35.5348 -1.3039 Td
[()]TJ
/F8 1 Tf
(.)Tj
/F14 1 Tf
0.9921 0 Td
[()-3()22()7()15()-13()12()]TJ
/F8 1 Tf
3.3392 0 Td
(subsp.)Tj
/F14 1 Tf
2.7212 0 Td
[( )15(\n)4()22()7()12()-4()]TJ
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2.7553 0 Td
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1.0374 0 Td
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2.5682 0 Td
(sense,)Tj
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1.4684 0 Td
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(of)Tj
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0.9297 0 Td
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3.8891 0 Td
([)Tj
0.83 0.64 0.02 0 k
(23)Tj
0 g
(].)Tj
2.0806 0 Td
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-35.5348 -1.2983 Td
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(1)Tj
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(\))Tj
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1.0205 0 Td
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(hundred)Tj
3.6737 0 Td
(plants.)Tj
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1.1055 0 Td
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2.0012 0 Td
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1.1736 0 Td
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3.1407 0 Td
(or)Tj
1.0999 0 Td
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4.5978 0 Td
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3.2031 0 Td
(of)Tj
1.0148 0 Td
(2)Tj
0.6973 0 Td
(to)Tj
1.0261 0 Td
(5)Tj
0.6917 0 Td
(plants.)Tj
2.8516 0 Td
(The)Tj
1.7858 0 Td
(number)Tj
3.4186 0 Td
(of)Tj
1.0148 0 Td
(sampled)Tj
3.5489 0 Td
(plants)Tj
2.6249 0 Td
(reflected)Tj
3.6453 0 Td
(population)Tj
4.5865 0 Td
(size)Tj
1.6894 0 Td
(estimated)Tj
4.0705 0 Td
(by)Tj
-35.8693 -1.2982 Td
(habitat)Tj
2.9876 0 Td
(survey,)Tj
3.0331 0 Td
(accordingly)Tj
4.8926 0 Td
(we)Tj
1.3266 0 Td
(sampled)Tj
3.5489 0 Td
(about)Tj
2.4945 0 Td
(every)Tj
2.3471 0 Td
(5)Tj
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Q
q
1 j
1 J
0 w
7.4999 0 0 6.6659 411.0803 520.6677 cm
BT
/F8 1 Tf
1 TL
-0.0053 Tc
0 0 Td
(th)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 419.4708 516.1889 cm
BT
/F8 1 Tf
1 TL
-0.004 Tc
0 0 Td
(-10)Tj
ET
Q
q
1 j
1 J
0 w
7.4999 0 0 6.6659 432.4535 520.6677 cm
BT
/F8 1 Tf
1 TL
-0.0053 Tc
0 0 Td
(th)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 440.844 516.1889 cm
BT
/F8 1 Tf
1 TL
-0.004 Tc
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(plants)Tj
2.6248 0 Td
(per)Tj
1.5307 0 Td
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1.4683 0 Td
(entire)Tj
2.5512 0 Td
(population)Tj
4.5921 0 Td
(area)Tj
1.8821 0 Td
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(Table)Tj
2.7836 0 Td
(1)Tj
0 g
(\).)Tj
1.2586 0 Td
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2.211 0 Td
(harvested)Tj
4.0422 0 Td
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2.5625 0 Td
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(plants)Tj
2.6249 0 Td
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(in)Tj
1.0318 0 Td
(silica)Tj
2.2223 0 Td
(gel)Tj
1.355 0 Td
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2.1033 0 Td
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1.7574 0 Td
(GPS)Tj
1.9729 0 Td
(positions)Tj
3.8551 0 Td
(were)Tj
2.1203 0 Td
(recorded)Tj
3.7984 0 Td
(\(by)Tj
1.5193 0 Td
(handheld)Tj
3.9855 0 Td
(Garmin)Tj
3.3732 0 Td
(receiver\))Tj
3.719 0 Td
(for)Tj
-35.5234 -1.3039 Td
(several)Tj
2.931 0 Td
(places)Tj
2.6248 0 Td
(at)Tj
0.9525 0 Td
(each)Tj
2.0296 0 Td
(locality.)Tj
ET
Q
q
1 j
1 J
0 w
11.9999 0 0 11.9999 200.0125 450.9921 cm
BT
/F15 1 Tf
1 TL
-0.0033 Tc
0 0 Td
(Description)Tj
5.159 0 Td
(of)Tj
1.063 0 Td
(habitat)Tj
ET
Q
q
1 j
1 J
0 w
10 0 0 10 200.0125 433.9842 cm
BT
/F8 1 Tf
1 TL
-0.004 Tc
0 0 Td
(The)Tj
1.7801 0 Td
(north-western)Tj
5.9357 0 Td
(Fertile)Tj
2.7779 0 Td
(Crescent)Tj
3.7304 0 Td
(in)Tj
1.0375 0 Td
(south-eastern)Tj
5.6522 0 Td
(Turkey)Tj
3.1068 0 Td
(is)Tj
0.8447 0 Td
(bordered)Tj
3.8834 0 Td
(by)Tj
1.1793 0 Td
(the)Tj
1.4683 0 Td
(Anti-Taurus)Tj
-31.3963 -1.2982 Td
(mountains)Tj
4.5297 0 Td
(to)Tj
1.0261 0 Td
(the)Tj
1.4684 0 Td
(Mesopotamian)Tj
6.2305 0 Td
(lowland,)Tj
3.6396 0 Td
(separating)Tj
4.3427 0 Td
(it)Tj
0.7823 0 Td
(from)Tj
2.2054 0 Td
(the)Tj
1.4683 0 Td
(central)Tj
2.9707 0 Td
(Anatolian)Tj
4.1839 0 Td
(plateau.)Tj
3.3279 0 Td
(It)Tj
-36.1755 -1.2983 Td
(is)Tj
0.8447 0 Td
(a)Tj
0.6576 0 Td
(region)Tj
2.8006 0 Td
(of)Tj
1.0148 0 Td
(rolling)Tj
2.8743 0 Td
(hills)Tj
1.8652 0 Td
(and)Tj
1.7291 0 Td
(a)Tj
0.6577 0 Td
(broad)Tj
2.5625 0 Td
(plateaus)Tj
3.4639 0 Td
(that)Tj
1.7801 0 Td
(extends)Tj
3.2655 0 Td
(into)Tj
1.8425 0 Td
(Syria.)Tj
2.4491 0 Td
(Eocene)Tj
3.1125 0 Td
(limestones)Tj
4.4787 0 Td
(with)Tj
-35.3988 -1.3039 Td
(small)Tj
2.3243 0 Td
(spots)Tj
2.2677 0 Td
(of)Tj
1.0148 0 Td
(basalt)Tj
2.5002 0 Td
(flows)Tj
2.3017 0 Td
(are)Tj
1.4513 0 Td
(characteristic)Tj
5.5446 0 Td
(of)Tj
1.0148 0 Td
(this)Tj
1.6724 0 Td
(area.)Tj
2.109 0 Td
(The)Tj
1.7858 0 Td
(limestone)Tj
4.1102 0 Td
(formation)Tj
4.2633 0 Td
(is)Tj
0.8447 0 Td
(dissected)Tj
-33.2048 -1.2983 Td
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1.1792 0 Td
(erosion)Tj
3.2088 0 Td
(and)Tj
1.7291 0 Td
(represents)Tj
4.3199 0 Td
(a)Tj
0.652 0 Td
(series)Tj
2.4265 0 Td
(of)Tj
1.0148 0 Td
(ridges)Tj
2.6362 0 Td
(\(~100)Tj
2.5681 0 Td
(m)Tj
1.0375 0 Td
(in)Tj
1.0375 0 Td
(height\))Tj
3.067 0 Td
(separated)Tj
4.0139 0 Td
(by)Tj
1.1792 0 Td
(wadis)Tj
2.4945 0 Td
(\(river)Tj
2.4491 0 Td
(val-)Tj
-35.0133 -1.2982 Td
(leys\).)Tj
2.2677 0 Td
(Quaternary)Tj
4.8302 0 Td
(sediments)Tj
4.2519 0 Td
(consist)Tj
2.9877 0 Td
(primarily)Tj
3.9685 0 Td
(of)Tj
1.0148 0 Td
(wash)Tj
2.228 0 Td
(and)Tj
1.7291 0 Td
(alluvial)Tj
3.0728 0 Td
(fan)Tj
1.491 0 Td
(deposits,)Tj
3.7077 0 Td
(as)Tj
1.0148 0 Td
(well)Tj
1.8255 0 Td
(as)Tj
1.0148 0 Td
(rela-)Tj
-35.4045 -1.3039 Td
(tively)Tj
2.364 0 Td
(thin)Tj
1.8595 0 Td
(\(1-m)Tj
2.2054 0 Td
(thick\))Tj
2.5738 0 Td
(and)Tj
1.7292 0 Td
(sporadic)Tj
3.6339 0 Td
(loamy)Tj
2.6816 0 Td
(slope)Tj
2.2734 0 Td
(deposits.)Tj
3.7077 0 Td
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1.1055 0 Td
(Sanliurfa,)Tj
4.0422 0 Td
(the)Tj
1.4683 0 Td
(average)Tj
3.2258 0 Td
(annual)Tj
-32.8703 -1.2983 Td
(temperature)Tj
5.125 0 Td
(is)Tj
0.8503 0 Td
(18.1C)Tj
2.9254 0 Td
(and)Tj
1.7348 0 Td
(average)Tj
3.2258 0 Td
(annual)Tj
2.9593 0 Td
(precipitation)Tj
5.3575 0 Td
(is)Tj
0.8504 0 Td
(447)Tj
1.6441 0 Td
(mm.)Tj
2.0862 0 Td
(The)Tj
1.7802 0 Td
(region)Tj
2.8063 0 Td
(belongs)Tj
3.2768 0 Td
(to)Tj
-34.6221 -1.2983 Td
(warm)Tj
2.5284 0 Td
(Mediterranean)Tj
6.1852 0 Td
(climate)Tj
3.1294 0 Td
(\(Csa\))Tj
2.3584 0 Td
(of)Tj
1.0148 0 Td
(Ko)Tj
0 Tc
0.737 0 Td
()Tj
-0.004 Tc
0.4422 0 Td
(ppen)Tj
2.2281 0 Td
(climate)Tj
3.1351 0 Td
(types)Tj
2.2733 0 Td
(and)Tj
1.7291 0 Td
(to)Tj
1.0262 0 Td
(Irano-Turanian)Tj
6.514 0 Td
(phytogeo-)Tj
-33.3012 -1.2982 Td
(graphical)Tj
3.9061 0 Td
(region.)Tj
3.0273 0 Td
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2.5796 0 Td
(are)Tj
1.4456 0 Td
(hot)Tj
1.5534 0 Td
(and)Tj
1.7291 0 Td
(dry)Tj
1.5761 0 Td
(summers)Tj
3.9004 0 Td
(\(mean)Tj
2.7893 0 Td
(July)Tj
1.7801 0 Td
(temperature)Tj
5.125 0 Td
(is)Tj
0.8504 0 Td
(31.6C;)Tj
3.1521 0 Td
(precipita-)Tj
-33.4145 -1.304 Td
(tion)Tj
1.8425 0 Td
(is)Tj
0.8447 0 Td
(2)Tj
0.6973 0 Td
(mm\))Tj
2.1997 0 Td
(and)Tj
1.7291 0 Td
(mild)Tj
2.0806 0 Td
(and)Tj
1.7291 0 Td
(comparatively)Tj
5.8734 0 Td
(humid)Tj
2.8913 0 Td
(winters)Tj
3.1691 0 Td
(\(mean)Tj
2.7893 0 Td
(January)Tj
3.3165 0 Td
(temperature)Tj
5.1307 0 Td
(is)Tj
-34.2933 -1.2982 Td
(5.0C,)Tj
2.6758 0 Td
(precipitation)Tj
5.3632 0 Td
(is)Tj
0.8447 0 Td
(119)Tj
1.6497 0 Td
(mm;)Tj
0.83 0.64 0.02 0 k
2.0806 0 Td
(www.globalbioclimatics)Tj
9.5074 0 Td
(.org)Tj
0 g
(\))Tj
2.126 0 Td
(i.e.)Tj
1.3606 0 Td
(semi-humid)Tj
5.1136 0 Td
(steppe)Tj
2.761 0 Td
(climate.)Tj
-33.4826 -1.2983 Td
(The)Tj
1.7801 0 Td
(vegetation)Tj
4.3426 0 Td
(comprises)Tj
4.269 0 Td
(\(semi-\)deciduous)Tj
7.217 0 Td
(oak)Tj
1.6554 0 Td
(wood-pasture)Tj
5.7316 0 Td
(dominated)Tj
4.5694 0 Td
(by)Tj
/F14 1 Tf
1.1849 0 Td
[()]TJ
/F8 1 Tf
(.)Tj
/F14 1 Tf
1.1509 0 Td
[()12()11()9( )7(\r)-4()11()11()-9()4()]TJ
/F8 1 Tf
-31.9009 -1.3039 Td
(subsp.)Tj
/F14 1 Tf
2.7155 0 Td
[()4()11()-4()10()10()4( )2()-9()]TJ
/F8 1 Tf
3.3789 0 Td
(or)Tj
/F14 1 Tf
1.0942 0 Td
[()]TJ
/F8 1 Tf
(.)Tj
/F14 1 Tf
1.1508 0 Td
[()20()13()11()12()]TJ
/F8 1 Tf
2.4095 0 Td
(subsp.)Tj
/F14 1 Tf
2.7212 0 Td
[()4( )7()12()12()6(\r)1()4(\n)12()12()-36(\n)4()11()20()]TJ
/F8 1 Tf
5.8563 0 Td
(\(K.Koch\))Tj
3.9345 0 Td
(Menitsky)Tj
3.9628 0 Td
(on)Tj
1.2699 0 Td
(neutral)Tj
3.0728 0 Td
(or)Tj
1.0998 0 Td
(alkaline)Tj
-32.6662 -1.2983 Td
(soils)Tj
1.9615 0 Td
(with)Tj
1.9956 0 Td
(relatively)Tj
3.8381 0 Td
(high)Tj
2.0126 0 Td
(organic)Tj
3.2371 0 Td
(content)Tj
3.2655 0 Td
([)Tj
0.83 0.64 0.02 0 k
(24)Tj
0 g
(].)Tj
2.0749 0 Td
(The)Tj
1.7859 0 Td
(typical)Tj
2.8573 0 Td
(habitat)Tj
2.9877 0 Td
(was)Tj
1.7007 0 Td
(ungrazed)Tj
3.9288 0 Td
(or)Tj
1.0999 0 Td
(slightly)Tj
-32.7456 -1.2982 Td
(grazed)Tj
2.8516 0 Td
(rocky)Tj
2.4775 0 Td
(limestone)Tj
4.1102 0 Td
(ground)Tj
3.1691 0 Td
(with)Tj
1.9956 0 Td
(scattered)Tj
3.7813 0 Td
(small)Tj
2.3301 0 Td
(\(2-4m\))Tj
3.0217 0 Td
(oak)Tj
1.6554 0 Td
(trees)Tj
/F14 1 Tf
2.0977 0 Td
[()-4()4( )2()20(\r)1()-4()]TJ
3.4355 0 Td
[()10()]TJ
/F8 1 Tf
(.)Tj
1.2529 0 Td
(accompanied)Tj
-32.1786 -1.3039 Td
(by)Tj
/F14 1 Tf
1.1792 0 Td
[()2()-4()9()11()6(\r)1()4()]TJ
3.2768 0 Td
[()11( )2()20( )20()11()12()22()10()18()-4()]TJ
/F8 1 Tf
4.5978 0 Td
(L.,)Tj
/F14 1 Tf
1.2075 0 Td
[()-2()11()-20()3(\n)12()-4()]TJ
3.1918 0 Td
[()22()3( )15(\n)4(\n)4()16()6()]TJ
/F8 1 Tf
3.5036 0 Td
(L.,)Tj
/F14 1 Tf
1.2132 0 Td
[()13()20()22()11()6( )9()-8()-4()]TJ
4.1329 0 Td
[()6()11()6()2()-21()-13()6()]TJ
/F8 1 Tf
4.32 0 Td
(Jacq.,)Tj
/F14 1 Tf
2.3641 0 Td
[()-11()-13()-9()-4()]TJ
2.4207 0 Td
[(\r)7()11()18()18()12()18()-4()]TJ
/F8 1 Tf
4.2633 0 Td
(L.,)Tj
/F14 1 Tf
-35.6709 -1.2983 Td
[()30(\r)7( )2()]TJ
1.9955 0 Td
[(\r)7()16()15()4( )8()9()7()20( )]TJ
4.1046 0 Td
[()]TJ
/F8 1 Tf
(.,)Tj
/F14 1 Tf
1.1962 0 Td
[()-2( )2()20()22()11()11()18()4()]TJ
4.1272 0 Td
[()10()4(\n)12()4()14()4( )]TJ
/F8 1 Tf
2.7156 0 Td
(L.,)Tj
/F14 1 Tf
1.2075 0 Td
[()23()11(\n)12()12()12()-9()-4()]TJ
3.5036 0 Td
[()10()11()12()6(\r)13()18()-9()-4()9()7()]TJ
/F8 1 Tf
5.04 0 Td
(Mill.,)Tj
/F14 1 Tf
2.3244 0 Td
[()-2( )2()20(\r)1()-4()]TJ
2.5625 0 Td
[()10()4(\n)12()4()14()4()9()7()-9()12()]TJ
/F8 1 Tf
5.1023 0 Td
(L.)Tj
0.9808 0 Td
(The)Tj
-34.8602 -1.2983 Td
(undergrowth)Tj
5.4708 0 Td
(indicates)Tj
3.77 0 Td
(presence)Tj
3.7134 0 Td
(of)Tj
1.0148 0 Td
(gaps)Tj
1.9956 0 Td
(in)Tj
1.0374 0 Td
(the)Tj
1.4684 0 Td
(canopy)Tj
3.1067 0 Td
(and)Tj
1.7292 0 Td
(agro-silvopastoral)Tj
7.353 0 Td
(land)Tj
1.9786 0 Td
(use.)Tj
1.7574 0 Td
(It)Tj
0.8561 0 Td
(con-)Tj
-35.2514 -1.3039 Td
(sists)Tj
1.8651 0 Td
(of)Tj
1.0148 0 Td
(heliophilous)Tj
5.1477 0 Td
(plants)Tj
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